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2022年12月29日发(作者:迅网卡盟)

中国毛茛科翠雀属的分类学研究(五):秋翠雀花二新异名

袁琼;杨亲二

【摘要】通过标本室和野外观察,发现根据四川西南部越西标本描述的毛茛科凉

山翠雀花()和根据四川西南部木里标本描

述的光轴翠雀花()只是此前记载分布于四川木里、盐

源和云南西北部宁蒗一带的秋翠雀花(aleHand.-Mazz.)植株较高的类

型,故将二者均处理为秋翠雀花的异名。%DelphiniumliangshanenW.T.

(Ranunculaceae)weredescribed

respectivelyfromYuexiandMuliinsouthwesternSichuan,

herbariumandifeldobrvationshaveshownthattheyreprentnomore

aleHand.-Mazz.,aspeciespreviously

recordedtooccurinsouthwesternSichuan(Muli,Yanyuan)and

northwesternYunnan(Ninglang).Wethereforereducethemtothe

ale.

【期刊名称】《热带亚热带植物学报》

【年(卷),期】2015(000)003

【总页数】9页(P268-276)

【关键词】翠雀属;毛茛科;分类学

【作者】袁琼;杨亲二

【作者单位】中国科学院华南植物园,中国科学院植物资源保护与可持续利用重点

实验室,广州510650;中国科学院华南植物园,中国科学院植物资源保护与可持

续利用重点实验室,广州510650

【正文语种】中文

In1931,Handel-Mazzetti[1]describedDelphiniumautumnaleHand.-

Mazz.(Ranunculaceae)onthebasisofacollectionofhisown,Handel-

Mazzetti5495(E,WU;Fig.1:A,B),fromYanyuan,southwestern

Sichuan,ngtimethisspecieshadnotbeenwell

1962,

Wang[2],

thefirsttime,aleasanindependentspecies,but

1967,Munz[3]

includedthespeciesinhispaper“AsynopsisoftheAsianspeciesof

Delphinium,nsustricto”,mentioningthathehadonlyexaminedan

isotypekeptinE(Fig.1:B).ale,

.

,butthelattertwospeciesaresodistinctfromit

thatthecomparisonisnotveryhelpfulforthedeterminationofitsidentity

1979,withoutgivinganyreason,Wang[4]reducedD.

enHuthasa

variety,ale(Hand.-Mazz.)1983,Wang

[5]foundaspecimen,al.103(PE;Fig.1:C)from

NinglanginnorthwesternYunnan,anareaclolycontiguoustoYanyuan

insouthwesternSichuan,ale,tomatch

nthisspecimen

aleasreprentingprobablyatransition

flora

alewasmostclolysimilar

,aspecieswithintheformer

subction,intheinflorescenceandflowercharacters,butverysimilar

florumL.,twospecieswithinthelatter

subction,intheleavesfinelydisctedwiththeultimate

lobulesnarrowlylinearandinthesuborbicularlimbsofthestaminode

en

florumbytheleavesmostlybasal,thestemleavesabnt

(ifprentthenverysmall),theinflorescencefewer-flowered,the

flowerslarger,andthespuroftheupperpalrecurved(hedidnot

hamatum).

Asaresult,alewasadistinct

phora,andthusimmediatelyresurrected

eatmenthasbeenadoptedbyWang[6-7]and

WangandWarnock[8].

In1979,Wang[9]

onthebasisofacollection,.

26197(Fig.2),

,aspeciesdescribed

fromnorthwesternYunnanandyetimperfectlyknown(onlyknownfrom

thetypematerial).AftercomparingtheirtypespecimenswefoundD.

aleinallesntialxceptforthe

niumliangshanenappearstobe

ale.

In1987,Wang[10]describedanotherspecies,Delphinium

,onthebasisofasinglespecimen,.

n.(Fig.3:A),fromMuli,southwesternSichuan,whichis

clolycontiguoustoYanyuan,thetypelocalityofD.

om

Muliandalsowiththeverywidespread,florum.

hanen,ourcomparisonoftheirtype

achyumisidenticalwithD.

autumnaleinallesntialxceptfortheslightdifferencein

niumleiostachyumappearstobesomewhattallerthanthe

ale.

AfterexaminingthetypematerialandotherspecimensofDelphinium

autumnaleweagreewithWang[5]totreatitasanindependentspecies

vesareusuallyremotefrom

eachotheronthestem;theleafbladeisfinelydiscted,withthe

primarylobessoclotoeachotherthatthebasalsinusverynarrowand

theultimatelobulesnarrowlylinear;thestemisdenlystrigoatba,

butglabrousoronlyverysparlyhispidinmedianandupperparts;the

pedicelsareglabrous,

botanicalexpeditiontosouthwesternSichuanandnorthwesternYunnanin

2005,wefoundthespeciestobefairlycommoninMulianditsadjacent

ky,grassyslopeatKangwuliangzi(misspeltas

KangduoliangzibyWang[6])inMuli,thetypelocalityof

achyum,wediscoveredalargepopulationofD.

eighborhoodwe

discoveredanothertwolargepopulationsalsoconsistingofplantsof

enrockyandgrassyplaceinNinglang,

northwesternYunnan,ale

showninFigures1-6,aleissomewhatvariableinstatureand

intheprenceorabnceofstemleaves,andinthetwocharacters

achyumfallwithintherangeof

identthatwhenWang[5]in1983

alehedidnotfullyunderstand

thespeciesbecauhisunderstanding,asmentionedabove,wasbad

ononlyasinglespecimen.

Badontheaboveanalys,weconsideritjustifiableto

achyumtothe

ale.

Taxonomictreatment

DelphiniumautumnaleHand.-.7:276.1931;W.T.

.10:274.1962;.48:503.1967;

.5:161.1983;an

Mount.1:489.1993;.11:131.2000;W.T.

Wang&6:260.2001.——envar.

autumnale(Hand.-Mazz.)risSin.27:

:wan(=Sichuan)austro-occid.:inmontis

Hwang-liangd,interoppidumYenyuenetcastellumKwapiregione

frigidetemperata,inpascuitadbambuta,3600-3900m,Oct.5,

1914,-Mazzetti5495(holotype,WU!;isotype,E!).

risSin.27:

615et425.1979;&6:.

:n:Yuexi,BijiShan,grassyslope,3000m,

Oct.4,1960,.26197(holotype,NAS!;

isotype,SM!).

.,Harbin7(2):96.1987;et

anMount.1:490.1993;&WarnockinFl.

China6::n:Muli,Kangwuliangzi,

alpinescrub,3600m,Aug.21,1983,.n.(holotype,

PE!).

n:Muli,Qinghai-Xizang

Exped.13215(PE),3373(PE),&337

(PE),339(PE),341(PE);Yanyuan,&510

(PE),al.3369(IBSC).Yunnan:Ninglang,

al.3340(IBSC),103(PE),&361

(PE),363(PE).

ralaspectandgeographicaldistribution,

DelphiniumautumnalemaybemostclolyrelatedtoD.

,aspeciesdescribedfromMuliin

southwesternSichuanandyetimperfectlyknownbecauofthe

ocentroides,thetwo

primarylaterallobesoftheleafbladearelessclotoeachotherandthus

thebasalsinusoftheleafbladeappearstobebroader,andallthethree

primarylobesarelessdiscted,withtheultimatelobulesslightly

ruerelationshipneedsafurther

study.

niumautumnaleisdistributedin

southwesternSichuan(Muli,Yanyuan,Yuexi)andnorthwestern

Yunnan(Ninglang),China(Fig.7).Itusuallygrowsinscruboron

grassy,rockyslopesataltitudesof3000-3900m.

AcknowledgmentsWearegratefultothecuratorsofE,IBSC,KUN,

NAS,PE,SMandWUforthepermissiontoutheirscannedimagesof

-minLiforthe

preparationofillustrations.

[1]ulaceae[M]//SymbolaeSinicae,VII.

:VerlagvonJuliusSpringer,1931:265-321.

[2]eskijobzorrodaDelphiniumizljutikovychflorykitaja

[J].ActaBotSin,1962,10(2):137-165.(inChine)

[3]sisoftheAsianspeciesofDelphinium,nsu

stricto[J].JArnArb,1967,48(4):476-545.

[4]niumL.[M]//FloraReipublicaePopularisSinicae,

g:SciencePress,1979:326-462.(inChine)

[5]edeRanunculaceissinensibus(VI)[J].Acta

BotYunnan,1983,5(2):153-163.(inChine)

[6]niumL.[M]//VascularPlantsoftheHengduan

Mountains,g:SciencePress,1993:474-493.(inChine)

[7]niumL.[M]//FloraYunnanica,Tomus11.

Beijing:SciencePress,2000:108-148.(inChine)

[8]WangWT,niumL.[M]//FloraofChina,Vol.

g:SciencePress&:MissouriBotanicalGardenPress,

2001:223-274.

[9]a[M]//FloraReipublicaePopularisSinicae,

g:SciencePress,1979:603-621.(inChine)

[10]edeRanunculaceissinensibus(XII)[J].Bull

BotRes(Harbin),1987,7(2):95-114.(inChine)

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